MD from the University of Valladolid (1989). PhD at the Institute of Biochemistry, CSIC-Universidad Complutense, Madrid (1994). Postdoctoral research fellow in Germany at the Max-Planck Institute for Biochemistry, Martinsried (1994-1997) and at the University of Cologne (1997-1998). Scientific assistant and Group leader at the Center for Biochemistry, Medical Faculty, University of Cologne (1998-2007). From 2007, Senior Lecturer in Biomedical Sciences at HYMS.